Monster Yamaha's Cal Crtuchlow recovered from early problems with the softer front Bridgestone tyre to set the fastest lap of the MotoGP weekend at Brno so far and topped this morning's third by a little less than two-tenths of a second.
The British rider, who complained of setup woes yesterday, switched to the harder option front with 18 minutes left and immediately jumped into the top four before going quickest with his final flying lap of 1'56.186.
Second-place man Jorge Lorenzo, who had led the session with a 1'56.332 set on his his sixth lap of 19, is 0.166s behind his fellow Yamaha teamster with Repsol Honda's Marc Marquez spoiling the Yamaha party in third place.
The MotoGP rookie was just able to stay in front of nine-time world champion Valentino Rossi in fourth and The Doctor is just a tenth down on the Spaniard after recording a 1'56.486, set with six minutes of the session left to run.
Dani Pedrosa is in fifth and half a second down on Crutchlow while Alvaro Bautista put in one quick lap to haul himself from the bottom end of the top ten to sixth and demote Bradley Smith to seventh place.
The young Briton was quicker than his Monster Yamaha team-mate early doors, putting in a 1'56.956 on lap seven of 18 which was good enough for fourth but he slipped down the order while concentrating on a race run.
Yesterday's fastest man Sefan Bradl couldn't muster anything like his time from free practice two and remaining in the 1'57s bracket to finish eighth. Andrea Dovizioso and Nicky Hayden round out the entries to qualifying two this afternoon.
Aleix Espargaro ended as top CRT runner with Hector Barbera and Colin Edwards behind him whike Michael Laverty has lost some of the ground he made up yesterday, slipping to 24th.